[1] Univ Alberta, Dept Biol Sci, Edmonton, AB, Canada
[2] CUNY Queens Coll, Dept Biol, Flushing, NY 11367 USA
[3] CUNY Queens Coll, Sch Earth & Environm Sci, Flushing, NY 11367 USA
[4] Xiamen Univ, Inst Marine Microbes & Ecospheres, Xiamen, Peoples R China
[5] Xiamen Univ, State Key Lab Marine Environm Sci, Xiamen, Peoples R China