[1] Museum Natl Hist Nat, Dept Systemat & Evolut, UMR Origine Struct & Evolut Biodiversite 7205, F-75231 Paris 05, France
[2] Univ Paris 11, UPS, CNRS, Lab Ecol Systemat & Evolut,CNRS,UMR 8079,AgroPari, F-91405 Orsay, France
[3] Univ Paris Diderot, Sorbonne Paris Cite, F-75475 Paris, France
[4] Museum Natl Hist Nat, Dept Hist Terre, UMR Paleobiodiversite Lignees & Communautes Anim, F-75231 Paris 05, France