[1] Univ Geneva, Inst Genet & Genom Geneva IGE3, Dept Genet & Evolut, CH-1211 Geneva 04, Switzerland
[2] Natl Inst Hlth, Dept Cell Biol & Neurosci, I-00161 Rome, Italy
[3] NOVA Univ Lisbon, CEDOC, NOVA Med Sch, P-1169056 Lisbon, Portugal
[4] Inst Pasteur, CNRS, Dept Dev & Stem Cell Biol, Stem Cells & Dev Unit,UMR 3738, F-75015 Paris, France