[1] Univ Penn, Dept Med, Perelman Sch Med, Philadelphia, PA 19104 USA
[2] Univ Penn, Dept Cell & Dev Biol, Perelman Sch Med, Philadelphia, PA 19104 USA
[3] Thomas Jefferson Univ, Sch Med, Dept Canc Biol, Philadelphia, PA 19107 USA
[4] Thomas Jefferson Univ, Sch Med, Dept Med, Philadelphia, PA 19107 USA