[1] Univ N Carolina, Dept Biol, Chapel Hill, NC 27599 USA
[2] Galapagos Sci Ctr, San Cristobal Isl, Galapagos Archi, Ecuador
[3] Univ Florida, Dept Biol, Gainesville, FL 32611 USA
[4] Univ Hamburg, Bioctr Grindel, D-20146 Hamburg, Germany
[5] Univ Hamburg, Zool Museum, D-20146 Hamburg, Germany
[6] Univ Barcelona, Dept Ecol, E-08028 Barcelona, Spain
[7] Univ Miami, Leonard & Jayne Abess Ctr Ecosyst Sci & Policy, Coral Gables, FL 33124 USA
[8] Univ Miami, RJ Dunlap Marine Conservat Program, Miami, FL 33149 USA
[9] Univ British Columbia, Dept Zool, Vancouver, BC V6T 1Z4, Canada