[1] Univ Chinese Acad Sci, Beijing 100049, Peoples R China
[2] Chinese Acad Sci, Northwest Inst Plateau Biol, Xining 810008, Peoples R China
[3] Chinese Acad Sci, Qinghai Inst Salt Lakes, CAS Key Lab Salt Lake Resources & Chenistry, Xining 810008, Peoples R China
[4] Qinghai Normal Univ, Sch Life & Geog Sci, Key Lab Qinghai Tibetan Plateau Environm & Resour, Xining 810008, Peoples R China