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South China Agr Univ, Key Lab Nat Pesticide & Chem Biol, Key Lab Crop Integrated Pest Management South Chi, Minist Agr,Minist Educ, Guangzhou 510642, Guangdong, Peoples R China South China Agr Univ, Key Lab Nat Pesticide & Chem Biol, Key Lab Crop Integrated Pest Management South Chi, Minist Agr,Minist Educ, Guangzhou 510642, Guangdong, Peoples R China