[1] Univ Zambia, Sch Vet Med, Dept Biomed Sci, Lusaka 10101, Zambia
[2] Univ Zambia, Hokudai Ctr Zoonosis Control Zambia, Lusaka 10101, Zambia
[3] Dept Med Control, Zambia Med Regulatory Author, Lusaka 10101, Zambia
[4] Univ Zambia, Sch Vet Med, Dept Dis Control, Lusaka 10101, Zambia
[5] Hokkaido Univ, Div Bioresources, Int Inst Zoonosis Control, N20 W10,Kita ku, Sapporo 0010020, Japan
[6] Hokkaido Univ, Int Inst Zoonosis Control, Div Int Res Promot, N20 W10,Kita ku, Sapporo 0010020, Japan
[7] Hokkaido Univ, Int Inst Zoonosis Control, Div Infect & Immun, N20 W10,Kita ku, Sapporo 0010020, Japan
[8] Hokkaido Univ, Inst Vaccine Res & Dev HU IVReD, N21 W11,Kita ku, Sapporo 0010020, Japan
[9] Hokkaido Univ, Int Inst Zoonosis Control, Div Mol Pathobiol, N20 W10,Kita ku, Sapporo 0010020, Japan
[10] Univ Zambia, Sch Vet Med, Dept Para Clin Studies, Lusaka 10101, Zambia
[11] Univ Zambia, Africa Ctr Excellence Infect Dis Humans & Anim, Sch Vet Med, Lusaka 10101, Zambia