[1] Spanish Res Council CSIC, Natl Inst Agr & Food Res & Technol INIA, Fish Immunol & Pathol Grp, Anim Hlth Res Ctr CISA,Anim Hlth Res Ctr CISA, Madrid, Spain
[2] Univ Porto, Ctr Interdisciplinar Invest Marinha & Ambiental C, Ave Gen Norton Matos S-N, P-4450208 Matosinhos, Portugal
[3] Univ Porto, Inst Ciencias Biomed Abel Salazar ICBAS, Rua Jorge Viterbo Ferreira 228, P-4050313 Porto, Portugal