[1] Sun Yat Sen Univ, Sch Marine Sci, Zhuhai 519082, Peoples R China
[2] Chinese Acad Sci, South China Sea Inst Oceanol, State Key Lab Trop Oceanog, Guangzhou 510301, Peoples R China
[3] Southern Marine Sci & Engn Guangdong Lab Zhuhai, Zhuhai 519080, Peoples R China
[4] Chinese Acad Sci, Inst Deep Sea Sci & Engn, Sanya 572000, Peoples R China