[1] Univ Oxford, Wellcome Ctr Human Genet, Nuffield Dept Med, Roosevelt Dr Headington, Oxford, England
[2] Univ Oxford, Dept Paediat, Oxford Vaccine Grp, Churchill Hospital Old Rd Headington, Oxford, England
[3] Univ Oxford, Chinese Acad Med Sci CAMS, Nuffield Dept Med, Oxford Inst, Old Rd Campus Headington, Oxford, England
[4] Univ Oxford, Wellcome Ctr Human Genet, Oxford, England