[1] Univ Cambridge, MRC Biostat Unit, Cambridge, England
[2] MRC UVRI & LSHTM, African Computat Genom TACG Res Grp, Entebbe, Uganda
[3] Univ Witwatersrand, Fac Hlth Sci, Sydney Brenner Inst Mol Biosci, Johannesburg, South Africa
[4] Univ Witwatersrand, Fac Hlth Sci, Dept Pediat, MRC Wits Dev Pathways Hlth Res Unit, Johannesburg, South Africa
[5] London Sch Hyg & Trop Med, Dept Noncommunicable Dis Epidemiol, London, England
[6] Univ Exeter, Exeter Ctr Excellence Diabet Res EXCEED, Sch Med, Exeter, Devon, England
[7] Univ Manchester, Ctr Genet & Genom Versus Arthrit, Ctr Musculoskeletal Res, Manchester, Lancs, England