[1] Univ Rey Juan Carlos, Dept Biol & Geol, Fis & Quim Inorgan, Madrid, Spain
[2] Univ Barcelona UB, Dept Biol Evolut Ecol & Ciencies Ambientals, Fac Biol, Barcelona, Spain
[3] Univ Barcelona UB, Inst Recerca Biodiversitat IRBio, Barcelona, Spain
[4] Oregon State Univ, Dept Forest Ecosyst & Soc, Corvallis, OR USA