[1] Univ Fed Sao Carlos, Lab Estudos Subterraneos, Sao Carlos, SP, Brazil
[2] Univ Cartagena, Grp Invest Biol Descript & Aplicada, Programa Biol, Campus San Pablo, Cartagena De Indias, Colombia
[3] Univ Cartagena, Grp Invest Hidrobiol, Programa Biol, Campus San Pablo, Cartagena De Indias, Colombia
[4] Univ Guajira, Fac Ingn, Grp Invest Terr Semiaridos Caribe, La Guajira, Colombia
[5] Univ Cyprus, Dept Biol Sci, Nicosia, Cyprus
[6] CNR, Ist Ric Ecosistemi Terrestri, Florence, Italy
[7] Museo Storia Nat, Sez Zool La Specola, Florence, Italy
[8] Inst Invest Recursos Biol Alexander von Humboldt, Secc Entomol, Colecc Biol, Villa De Leyva, Boyaca, Colombia