[1] Univ Concepcion, Fac Ciencias Nat & Oceanog, Dept Zool, Casilla 160-C, Concepcion, Chile
[2] Univ La Serena, Fac Ciencias, Dept Biol, Lab Entomol Ecol LEULS, Casilla 554, La Serena, Chile
[3] Inst Ecol & Biodiversidad IEB, Santiago, Chile
[4] Univ Concepcion, Fac Ciencias Nat & Oceanog, Dept Bot, Casilla 160-C, Concepcion, Chile