Lower Permian (Late Kungurian) conodonts from the Sibumasu Terrane, Malaysia: paleoecological, paleobiogeographical and tectonic implications

Late Kungurian (Lower Permian) conodonts are described from the Kanthan Limestone, Perak, Peninsular Malaysia and for the first time from the Sibumasu Terrane of the Malay Peninsula. The co-occurrence of Gullodus duani, Gullodus hemicircularis, Gullodus sicilianus, Mesogondolella lamberti and Mesogondolella siciliensis represent the Mesogondolella lamberti International Conodont Zone and the broadly equivalent Mesogondolella siciliensis Regional Conodont Zone. A small fault-bounded basal Pennsylvanian (basal Bashkirian) conodont fauna including Gnathodus girtyi simplex and Declinognathodus inaequalis is also reported. The late Kungurian conodonts from the Kanthan Limestone were deposited in a relatively deep-water environment on the northern passive margin of the Sibumasu Terrane of the eastern Cimmerian Continent located at c. 35(o)S latitude. Biogeographically, the fauna represents the southern peri-Gondwana Cool Water Province which is consistent with its palaeogeographic location. A new scheme, utilizing characteristics of P1 elements, including position of the 1st denticle, location of 2nd and 3rd denticles, platform shape, platform cross-section, denticle shape in cross-section, and lateral denticle development is proposed for distinguishing between species of the hindeodid genera Gullodus, Hindeodus and Isarcicella. The late Kungurian fauna from the Kanthan Limestone represents the southern peri-Gondwana Cool Water Province supporting palaeogeographic reconstructions placing the Sibumasu Terrane in moderate southern palaeolatitudes in the Kungurian.