[1] Yunnan Univ, Sch Ecol & Environm Sci, Kunming 650500, Yunnan, Peoples R China
[2] Yunnan Univ, Sch Life Sci, Kunming 650500, Yunnan, Peoples R China
[3] Chinese Acad Sci, Chengdu Inst Biol, CAS Key Lab Mt Ecol Restorat & Bioresource Utiliz, Chengdu, Peoples R China
[4] Chinese Acad Sci, Chengdu Inst Biol, Ecol Restorat & Biodivers Conservat Key Lab Sichu, Chengdu, Peoples R China
[5] Chinese Acad Sci, Kunming Inst Bot, CAS Key Lab Plant Divers & Biogeog East Asia, Kunming 650201, Yunnan, Peoples R China