[1] Univ Fed Para, Ctr Estudos Avancados Biodiversidade, Lab Genet & Biol Celular, Inst Ciencias Biol, BR-66075110 Belem, Para, Brazil
[2] Sao Paulo State Univ, Inst Biosci Botucatu, Dept Struct & Funct Biol, UNESP, BR-18618689 Botucatu, SP, Brazil
[3] Univ Fed Para, Lab Oleos Amazonia, BR-66075110 Belem, Para, Brazil
[4] Inst Biol & Conservacao Mamiferos Aquat Amazonia B, BR-66077830 Belem, Para, Brazil