[1] Minist Nat Resources, Inst Oceanog 1, Marine Bioresource & Environm Res Ctr, Key Lab Marine Ecoenvironm Sci & Technol, Qingdao 266061, Peoples R China
[2] China Univ Geosci, Sch Ocean Sci, Beijing 100083, Peoples R China
[3] Qingdao Natl Lab Marine Sci & Technol, Lab Marine Ecol & Environm Sci, Qingdao 266071, Peoples R China