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Univ Fed Pernambuco, Ctr Biociencias, Dept Zool, Pos Grad Biol Anim, Ave Nelson Chaves S-N, BR-50373970 Recife, PE, Brazil Univ Fed Pernambuco, Ctr Biociencias, Dept Zool, Pos Grad Biol Anim, Ave Nelson Chaves S-N, BR-50373970 Recife, PE, Brazil