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Huazhong Univ Sci & Technol, Key Lab Mol Biophys, Minist Educ,Dept Bioinformat & Syst Biol,Coll Lif, Hubei Key Lab Bioinformat & Mol Imaging,Ctr AI Bi, Wuhan, Peoples R China Chinese Acad Sci, Inst Hydrobiol, Natl Aquat Biol Resource Ctr, Wuhan, Peoples R China