[1] Univ St Andrews, Ctr Res Ecol & Environm Modelling, St Andrews, Scotland
[2] Univ Oxford, Dept Biol, Wildlife Conservat Res Unit, Oxford, England
[3] Cornell Univ, Dept Nat Resources & Environm, Ithaca, NY USA
[4] NOAA Natl Marine Fisheries Serv, Northeast Fisheries Sci Ctr, Woods Hole, MA USA
[5] Univ Oxford, Dept Biol, Oxford, England
[6] Zambian Carnivore Programme, Mfuwe, Zambia