[1] Tech Univ Denmark, Natl Inst Aquat Resources DTU Aqua, North Sea Sci Pk, DK-9850 Hirtshals, Denmark
[2] Leibniz Inst Freshwater Ecol & Inland Fisheries, Dept Fish Biol Fisheries & Aquaculture, Berlin, Germany
[3] Univ Highlands & Isl, Shetland UHI, Scalloway Campus, Scalloway ZE1 0UN, Zetland, Scotland