[1] World Vegetable Ctr, Tainan 74151, Taiwan
[2] CIFOR ICRAF, Trees Res Theme, World Agroforestry, Nairobi 00100, Kenya
[3] Univ Abomey Calavi, Fac Agron Sci, Unit Genet Biotechnol & Seed Sci, 01 BP 526, Cotonou, Benin
[4] World Vegetable Ctr, Eastern & Southern Africa, Arusha, Tanzania
[5] Natl Taiwan Univ, Dept Agron, Taipei 10617, Taiwan
[6] Scotlands Rural Coll, Principals Res Grp, Edinburgh EH9 3JG, Midlothian, Scotland