[1] Okayama Univ Sci, Fac Vet Med, Lab Vet Pathol, Imabari, Ehime, Japan
[2] Kyoto Inst Nutr & Pathol Inc, Ujitawara, Kyoto, Japan
[3] Setsunan Univ, Fac Pharmaceut Sci, Lab Pathol, Hirakata, Osaka, Japan
[4] Tsushima Wildlife Conservat Ctr, Saozakikoen, Tsushima, Nagasaki, Japan
[5] Wildlife Management Off, West Branch, 3457-1 Arinocho Arino,Kita ku, Kobe, Hyogo 6511312, Japan