[1] Anhui Normal Univ, Coll Life Sci, Anhui Prov Key Lab Conservat & Exploitat Biol Reso, Wuhu 241000, Peoples R China
[2] Anhui Agr Univ, Sch Life Sci, Hefei 230036, Peoples R China
[3] Huangshan Noah Biodivers Inst, Huangshan 245000, Peoples R China
[4] Shanghai Ocean Univ, Shanghai Collaborat Innovat Aquat Anim Genet & Bre, Shanghai 201306, Peoples R China
[5] Nanning Municipal Publ Secur Bur, Nanning 530012, Peoples R China
[6] Rutgers State Univ, Dept Ecol Evolut & Nat Resources, Grad Program Ecol & Evolut, New Brunswick, NJ 08901 USA