[1] East Carolina Univ, Dept Coastal Studies, 850 NC 345, Wanchese, NC 27981 USA
[2] Univ Calif Santa Cruz, Dept Ocean Sci, 1156 High St, Santa Cruz, CA 95064 USA
[3] Univ Calif Santa Cruz, Inst Marine Sci, 115 McAllister Way, Santa Cruz, CA 95060 USA
[4] ERM, Siemensstr 9, D-63263 Neu Isenburg, Germany
[5] Nature Conservancy, Berlin, Germany
[6] Humboldt Univ, Berlin, Germany