[1] Inst Nacl Biodivers INABIO, Unidad Invest, Quito, Ecuador
[2] Fdn EcoMinga, Banos, Ecuador
[3] Fdn Oscar Efren Reyes, Dept Ambiente, Banos, Ecuador
[4] Univ San Francisco Quito USFQ, Colegio Ciencias Biol & Ambientales COCIBA, Lab Biol Evolut, Quito, Cumbaya, Ecuador
[5] Univ Complutense Madrid, Madrid, Spain
[6] Univ San Francisco Quito USFQ, Museo Zool, Inst IBIOTROP, Quito, Ecuador
[7] Univ San Francisco Quito USFQ, Colegio Ciencias Biol & Ambientales COCIBA, Lab Zool Terr, Quito, Ecuador