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Anqing Normal Univ, Sch Life Sci, Prov Key Lab Biodivers Study & Ecol Conservat Sou, Res Ctr Aquat Organism Conservat & Water Ecosyst, Anqing City 246011, Anhui, Peoples R China Anqing Normal Univ, Sch Life Sci, Prov Key Lab Biodivers Study & Ecol Conservat Sou, Res Ctr Aquat Organism Conservat & Water Ecosyst, Anqing City 246011, Anhui, Peoples R China