[1] Univ Cambridge, Dept Pathol, Cambridge, England
[2] Lyzeum Ltd, Cambridge, England
[3] Univ Edinburgh, Dept Pathol, Edinburgh, Midlothian, Scotland
[4] Western Gen Hosp, Dept Pathol, NHS Lothian, Edinburgh, Midlothian, Scotland
[5] Univ Hosp Birmingham NHS Trust, Heartlands Hosp, Dept Cellular Pathol, Birmingham, W Midlands, England
[6] Univ Cambridge, Dept Appl Math & Theoret Phys, Wilberforce Rd, Cambridge CB3 0WA, England
[7] Univ Hosp North Tees, North Tees & Hartlepool NHS Fdn Trust, Stockton On Tees, England