[1] Univ Tours, INRAE, CNRS, PRC, Nouzilly, France
[2] Univ Tours, INRAE, BOA, F-37380 Nouzilly, France
[3] Univ Tours, INRAE, ISP, UMR 1282, Nouzilly, France
[4] Univ Toulouse, INRAE INPT ESUPAT, GenPhySE, Castanet Tolosan, France
[5] INRAE, PEAT, Nouzilly, France
[6] INRAE, Elevage Alternat & Sante Monogastr UE1206, St Pierre Damilly, France