[1] Univ Laval, Dept Biol, Quebec City, PQ, Canada
[2] Univ Laval Canada, Takuvik Int Res Lab, LRI3376, Quebec City, PQ, Canada
[3] CNRS, Paris, France
[4] MIT, Dept Earth Atmospher & Planetary Sci, Cambridge, MA USA
[5] MIT, Ctr Global Change Sci, Cambridge, MA USA
[6] Inst Maurice Lamontagne Peches & Oceans Canada, Mont Joli, PQ, Canada
[7] Univ Quebec Rimouski UQAR, Inst Sci Mer Rimouski ISMER, Rimouski, PQ, Canada
[8] Univ Laval, Dept Biochem Microbiol & Bioinformat, Quebec City, PQ, Canada
[9] UBO, Lab Sci Environm Marin LEMAR, Inst Univ Europeen Mer IUEM,Technopole Brest Irois, CNRS,IRD,Ifremer, Plouzane, France