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Univ Liverpool, Inst Syst Mol & Integrat Biol, Ctr Metabol Res, Dept Biochem Cell & Syst Biol, Biosci Bldg,Crown St, Liverpool L69 7ZB, England Univ Liverpool, Inst Syst Mol & Integrat Biol, Ctr Metabol Res, Dept Biochem Cell & Syst Biol, Biosci Bldg,Crown St, Liverpool L69 7ZB, England