[1] Key Laboratory of the Ministry of Education for Protection and Utilization of Special Biological Resources in the Western, School of Life Science, Ningxia University, Ningxia, Yinchuan,750021, China
[2] College of Life Sciences, Gannan Normal University, Jiangxi, Ganzhou,341000, China
[3] Key Laboratory of Adaptation and Evolution of Plateau Biota (AEPB), Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Qinghai, Xining, China
[4] Guangdong Provincial Key Laboratory of Applied Botany, State Key Laboratory of Plant Diversity and Specialty Crops, South China Botanical Garden, Chinese Academy of Sciences, South China National Botanical Garden, Guangzhou,510650, China
[5] College of Agricultural Science, Ningxia University, Ningxia, Yinchuan,750021, China