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Univ Alberta, Dept Med, Div Gastroenterol,Nutr & Metab Res Grp, Cell & Mol Biol Collaborat Network Gastrointestin, Edmonton, AB T6G 2C2, Canada Univ Alberta, Dept Med, Div Gastroenterol,Nutr & Metab Res Grp, Cell & Mol Biol Collaborat Network Gastrointestin, Edmonton, AB T6G 2C2, Canada