[1] Bristol Palaeobiology Group, School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol,BS8 1TQ, United Kingdom
[2] State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing,210008, China
[3] CAS Center for Excellence in Life and Paleoenvironment, Nanjing,210008, China