[1] Sun Yat Sen Univ, Sch Marine Sci, Guangzhou 510275, Peoples R China
[2] Guangdong Prov Key Lab Marine Resources & Coastal, Guangzhou 510275, Peoples R China
[3] Minist Educ, Pearl River Estuary Marine Ecosyst Res Stn, Zhuhai 519082, Peoples R China
[4] Southern Marine Sci & Engn Guangdong Lab, Zhuhai, Guangdong, Peoples R China
[5] Beibu Gulf Univ, Coll Marine Sci, Guangxi Key Lab Marine Environm Change & Disaster, Qinzhou 535011, Peoples R China