[1] Univ Calgary, Dept Geosci, 2500S Univ Dr NW, Calgary, AB T2N 1N4, Canada
[2] US Geol Survey, Woods Hole Coastal & Marine Sci Ctr, 384 Woods Hole Rd, Woods Hole, MA 02543 USA
[3] US Geol Survey, Pacific Coastal & Marine Sci Ctr, 2885 Mission St, Santa Cruz, CA 95060 USA