[1] Natl Inst Fisheries Sci, Ocean Climate & Ecol Res Div, Busan 46083, South Korea
[2] Natl Inst Fisheries Sci, Marine Environm Res Div, Pusan 46083, South Korea
[3] Pusan Natl Univ, Dept Oceanog, Busan 46241, South Korea
[4] Pusan Natl Univ, Marine Sci Inst, Busan 46241, South Korea