[1] Univ Pretoria, Dept Zool & Entomol, Private Bag X20, ZA-0028 Hatfield, South Africa
[2] Univ Fed Mato Grosso, Lab Scarabaeoidol, Inst Biociencias, BR-78060900 Cuiaba, MT, Brazil
[3] Univ Pernambuco, Dept Ciencias Biol, Campus Petrolina, BR-56328900 Petrolina, PE, Brazil
[4] Univ Fed Parana, Lab Sistemat & Bioecol Coleoptera, Dept Zool, BR-81531980 Curitiba, PR, Brazil
[5] Univ Sao Paulo, Museu Zool, BR-04218970 Sao Paulo, SP, Brazil
[6] Univ Fed Alagoas, Lab Sistemat & Diversidade Artropodes, Inst Ciencias Biol & Saude, BR-57072900 Maceio, AL, Brazil
[7] Univ Fed Rural Pernambuco, Dept Agron, Lab Taxon Insetos, Fitossanidade, BR-52171970 Recife, PE, Brazil