[1] Yangtze Univ, Sch GeoSci, Wuhan 430100, Hubei, Peoples R China
[2] Chinese Acad Sci, Nanjing Inst Geol & Palaeontol, State Key Lab Palaeobiol & Stratig, 39 East Beijing Rd, Nanjing 210008, Peoples R China
[3] Univ Antilles, Sorbonne Univ, Museum Natl Hist Nat, Inst Systemat,Evolut,Biodiversite,CNRS,ISYEB,UMR 7, CP 50,57 rue Cuvier, F-75005 Paris, France
[4] Jiangsu Second Normal Univ, Inst Insect Resources & Biodivers, Sch Life Sci Chem & Chem Engn, Nanjing 210013, Peoples R China
[5] Room 1402,22,Lane 999,Zhenghe Rd, Shanghai 200438, Peoples R China
[6] Univ Chinese Acad Sci, Beijing 100049, Peoples R China