[1] Univ Alberta, Dept Med Microbiol & Immunol, Edmonton, AB, Canada
[2] NIDCR, Mucosal Immunol Sect, NIH, Bethesda, MD 20892 USA
[3] Univ Seville, Sch Med, Dept Med Biochem Mol Biol & Immunol, Seville, Spain
[4] Univ Seville, Virgen Rocio Univ Hosp, Virgen Macarena Univ Hosp, Inst Biomed Seville IBiS,CSIC, Seville, Spain