[1] Univ Tennessee, Dept Anim Sci, Knoxville, TN 37996 USA
[2] Univ Tennessee, Dept Biosyst Engn & Soil Sci, Knoxville, TN 37996 USA
[3] Univ Helsinki, Fac Agr & Forestry, Dept Agr Sci, Helsinki 00014, Finland
[4] Univ Helsinki, Fac Vet Med, Res Ctr Anim Welf, Dept Prod Anim Med, Helsinki 00014, Finland
[5] Univ Helsinki, Helsinki One Hlth, FI-00014 Helsinki, Finland