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Shenyang Agr Univ, Rice Res Inst, Minist Agr Educ & Liaoning Prov, Key Lab Northeast Rice Biol & Genet & Breeding,Ke, Shenyang 110866, Peoples R China Shenyang Agr Univ, Rice Res Inst, Minist Agr Educ & Liaoning Prov, Key Lab Northeast Rice Biol & Genet & Breeding,Ke, Shenyang 110866, Peoples R China