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South China Agr Univ, Coll Anim Sci, Guangdong Prov Key Lab Agroanim Genom & Mol Breedi, State Key Lab Swine & Poultry Breeding Ind, Guangzhou 510640, Peoples R China South China Agr Univ, Coll Anim Sci, Guangdong Prov Key Lab Agroanim Genom & Mol Breedi, State Key Lab Swine & Poultry Breeding Ind, Guangzhou 510640, Peoples R China