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Yonsei Univ, Brain Korea Project 21, Dept Oral Biol,Human Identificat Res Ctr, Div Anat & Dev Biol,Oral Sci Res Ctr,Coll Dent, Seoul 120752, South Korea Yonsei Univ, Brain Korea Project 21, Dept Oral Biol,Human Identificat Res Ctr, Div Anat & Dev Biol,Oral Sci Res Ctr,Coll Dent, Seoul 120752, South Korea