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- [4] Changed energy state and increased mitochondrial β-oxidation rate in liver of rats associated with lowered proton electrochemical potential and stimulated uncoupling protein 2 (UCP-2) expression -: Evidence for peroxisome proliferator-activated receptor-α independent induction of UCP-2 expression JOURNAL OF BIOLOGICAL CHEMISTRY, 2003, 278 (33) : 30525 - 30533