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CSIC, Museo Nacl Ciencias Nat, Dept Biodivers & Evolutionary Biol, Jose Gutierrez Abascal 2, E-28006 Madrid, Spain
Srnithonian Inst, Dept Invertebrate Zool, Natl Museum Nat Hist, 10th St & Constitut Ave NW, Washington, DC 20560 USA
Univ Magdalena, Grp Evolut Sistemat & Ecol Mol, Cl 32 22-08, Santa Marta, Colombia CSIC, Museo Nacl Ciencias Nat, Dept Biodivers & Evolutionary Biol, Jose Gutierrez Abascal 2, E-28006 Madrid, Spain