[1] Chinese Acad Sci, Kunming Inst Bot, CAS Key Lab Plant Divers & Biogeog East Asia, Kunming 650201, Yunnan, Peoples R China
[2] Nat Resources Bur Guidong Cty, Chenzhou 423000, Hunan, Peoples R China
[3] Univ Chinese Acad Sci, Beijing 100049, Peoples R China