[1] Smithsonian Inst, Dept Paleobiol, Natl Museum Nat Hist, Washington, DC 20560 USA
[2] Univ Hong Kong, Swire Inst Marine Sci, Sch Biol Sci, Area Ecol & Biodivers, Kadoorie Biol Sci Bldg,Pokfulam Rd, Hong Kong, Peoples R China
[3] Univ Hong Kong, Swire Inst Marine Sci, State Key Lab Marine Pollut, Kadoorie Biol Sci Bldg,Pokfulam Rd, Hong Kong, Peoples R China
[4] US Geol Survey, Florence Bascom Geosci Ctr, MS 926A,12201 Sunrise Valley Dr, Reston, VA 20192 USA